Page 186 of 491

185 Controls in detail
Automatic transmission
One-touch gearshifting 
Even with an automatic transmission you 
can change the gears manually when the 
gear selector lever is in positionD.
Downshifting
Briefly press the gear selector lever to 
the left in theD-direction.
The transmission will shift from the current 
gear to the next lower gear. This action 
simultaneously limits the gear range of the 
transmission (
page 186).Upshifting
Briefly press the gear selector lever to 
the right in theD+direction.
The transmission will shift from the current 
gear to the next higher gear as permitted 
by the shift program. This action simul-
taneously extends the gear range of the 
transmission.Canceling gear range limit
Press and hold the gear selector lever 
in theD+direction until
D reappears in 
the multifunction display.
The transmission will shift from the current 
gear range directly to gear rangeD.
Shifting into optimal gear range 
Press and hold the gear selector lever 
in theD-direction.
The transmission will automatically select 
the gear range suited for optimal 
acceleration and deceleration. This will 
involve shifting down one or more gears.
Warning!
G
On slippery road surfaces, never downshift 
in order to obtain braking action. This could 
result in drive wheel slip and reduced 
vehicle control. Your vehicle’s ABS will not 
prevent this type of loss of control.
iTo avoid overrevving the engine when 
the gear selector lever is moved to 
theD-direction, the transmission will 
not shift to a lower gear if the engine’s 
max. speed would be exceeded. 
     
        
        Page 187 of 491

186 Controls in detailAutomatic transmissionGear ranges
With the gear selector lever in positionD, 
you can limit the transmission’s gear range 
by pressing the gear selector lever to the 
left (D-), and reverse the gear range limit 
by pressing the gear selector lever to the 
right (D+).
The selected gear range appears in the 
multifunction display (
page 184). If you 
press on the accelerator when the engine 
has reached its rpm limit, the transmission 
will upshift beyond any gear range limit se-
lected.
Effect
ï
The transmission shifts through 
sixth gear only (applies to 
vehicles with 7-speed automatic 
transmission only).
î
The transmission shifts through 
fifth gear only (applies to 
vehicles with 7-speed automatic 
transmission only).
é
The transmission shifts through 
fourth gear only.
è
The transmission shifts through 
third gear only.
With this selection you can use 
the braking effect of the engine.
Effect
ç
The transmission shifts through 
second gear only.
Allows the use of engine’s 
braking power when driving:
on steep downgrades
in mountainous regions
under extreme operating 
conditions
æ
The transmission operates in 
first gear only.
For maximum use of engine’s 
braking effect on very steep or 
lengthy downgrades. 
     
        
        Page 188 of 491

187 Controls in detail
Automatic transmission
Gear selector lever position
Effect
ì
Park position 
Gear selector lever position 
when the vehicle is parked. Place 
gear selector lever in positionP 
only when vehicle is stopped. 
The park position is not intended 
to serve as a brake when the 
vehicle is parked. Rather, the 
driver should always set the 
parking brake in addition to 
placing the gear selector lever in 
positionP to secure the vehicle.
EffectThe SmartKey can only be 
removed from the starter switch 
with the gear selector lever in 
positionP. With the SmartKey 
removed, the gear selector lever 
is locked in positionP.
If the vehicle’s electrical system 
is malfunctioning, the gear 
selector lever could remain 
locked in positionP 
(page 405).
í
Reverse gear 
Place gear selector lever in 
positionR only when vehicle is 
stopped.
Effect
ë
Neutral
No power is transmitted from the 
engine to the drive axle. When 
the brakes are released, the 
vehicle can be moved freely 
(pushed or towed).
To avoid damage to the trans-
mission, never engageN while 
driving.
If the ESP is deactivated or 
malfunctioning:
Move gear selector lever toN 
only if the vehicle is in danger of 
skidding, e.g. on icy roads.
ê
Drive 
The transmission shifts 
automatically. All forward gears 
are available. 
     
        
        Page 189 of 491

188 Controls in detailAutomatic transmission
Automatic shift program
The program mode selector switch is 
located on the lower part of the center 
console.
1Program mode selector switch
C Comfort For comfort driving
S Sport For standard driving
The current gear selector lever position 
and the selected program mode (
C/S) are 
indicated in the multifunction display 
(
page 184).
!Coasting the vehicle, or driving for any 
other reason with gear selector lever 
inN can result in transmission damage 
that is not covered by the 
Mercedes-Benz Limited Warranty.Warning!
G
Getting out of your vehicle with the gear 
selector lever not fully engaged in positionP 
is dangerous. Also, positionP alone is not 
intended to or capable of preventing your 
vehicle from moving, possibly hitting people 
or objects.
Always set the parking brake in addition to 
shifting to positionP (
page 56).
When parked on an incline, turn the front 
wheels towards the road curb.
Do not park this vehicle in areas where com-
bustible materials such as grass, hey or 
leaves can come into contact with the hot 
exhaust system, as these materials could be 
ignited and cause a vehicle fire.
Warning!
G
When leaving the vehicle, always remove the 
SmartKey or the SmartKey with 
KEYLESS-GO* from the starter switch, take 
it with you, and lock the vehicle. Do not 
leave children unattended in the vehicle, or 
w i t h  a c c e s s  t o  a n  u n l o c k e d  v e h i c l e .  C h i l d r e n  
could move the gear selector lever from 
positionP, which could result in an accident 
and/or serious personal injury. 
     
        
        Page 190 of 491

189 Controls in detail
Automatic transmission
Press program mode selector 
switch1 repeatedly until the letter of 
the desired shift program appears in 
the multifunction display.
SelectCfor comfort driving:
The vehicle starts out in second 
gear (both forward and reverse) for 
gentler starts. This does not apply if 
full throttle is applied or gear 
range1 is selected.
Traction and driving stability are 
improved on icy roads.
Upshifts occur earlier even when 
you give more gas. The engine then 
operates at lower rpms and the 
wheels are less likely to spin.
Driving tips
Accelerator position
Your driving style influences the 
transmission’s shifting behavior:
Less throttle Earlier upshifting
More throttle  Later upshifting
Kickdown 
Use kickdown when you want maximum 
acceleration.
Press the accelerator past the point of 
resistance.
The transmission shifts into a lower 
gear.
Ease on the accelerator when you have 
reached the desired speed. 
The transmission shifts up again.
!Never change the program mode when 
the gear selector lever is out of 
positionP. This could result in a 
change of driving characteristics for 
which you may not be prepared.iThe last selected program 
mode (CorS) is switched on when the 
engine is restarted. 
     
        
        Page 191 of 491

190 Controls in detailAutomatic transmissionStopping
When you stop briefly, e.g. at traffic lights:
Leave the transmission in gear.
Hold the vehicle with the brake.
When you stop longer with the engine 
idling or on an uphill gradient:
Move the gear selector lever to 
positionP.
Set the parking brake.Maneuvering
When you maneuver in tight areas, 
e.g. when pulling into a parking space:
Control the vehicle speed by gradually 
releasing the brakes.
Accelerate gently.
Never abruptly step on the accelerator.
Working on the vehicleEmergency operation 
(Limp Home Mode)
If vehicle acceleration worsens or the 
transmission no longer shifts, the trans-
mission is most likely operating in limp 
home (emergency operation) mode. In this 
mode only second gear and reverse gear 
can be activated.
Stop the vehicle.
Move gear selector lever toP.
Turn off the engine.
Wait at least ten seconds before 
restarting.
Restart the engine.
Move gear selector lever to positionD 
(for second gear) orR.
Have the transmission checked at an 
authorized Mercedes-Benz Center as 
soon as possible.
Warning!
G
When working on the vehicle, set the 
parking brake and move gear selector lever 
to positionP. Otherwise the vehicle could 
roll away. 
     
        
        Page 230 of 491

229 Controls in detail
Driving systems
1Set current or higher speed
2Set current or lower speed
3Cancel cruise control
4Resume at previously set speedSetting current speed
Accelerate or decelerate to the desired 
speed.
Briefly lift1 or depress2 the cruise 
control lever.
The current speed is set.
Remove your foot from the accelerator 
pedal.
Cruise control is activated.
The selected speed appears in the multi-
function display for approximately five sec-
onds, and the corresponding speedometer 
segments from the selected speed to the 
vehicle maximum speed are illuminated.
iOn uphill or downhill grades, cruise 
control may not be able to maintain the 
set speed. Once the grade eases, the 
set speed will be resumed.
E 320, E 500:
On downhill grades, the cruise control 
will hold the set speed with active brak-
ing action.
In addition, on longer downhill grades 
the automatic transmission will auto-
matically downshift.
E 320 4MATIC, E 500 4MATIC:
If the engine’s braking power does not 
brake the vehicle sufficiently on down-
hill grades, the automatic transmission
will automatically downshift. 
     
        
        Page 232 of 491

231 Controls in detail
Driving systems
Setting a higher speed
Lift cruise control lever to position1 
and hold it up until the desired speed is 
reached.
Release cruise control lever.
The new speed is set.
Setting a lower speed
Depress cruise control lever to 
position2 and hold it down until the 
desired speed is reached.
Release cruise control lever.
The new speed is set.Fine adjustment in 1 mph 
(Canada: 1 km / h) increments
Faster
Briefly tip cruise control lever in the di-
rection of arrow1.
Slower
Briefly tip cruise control lever in the di-
rection of arrow2.Setting to last stored speed 
(“Resume” function)
Briefly push cruise control lever to 
position4.
The cruise control resume the last set 
speed.
Remove your foot from the accelerator 
pedal.
The selected speed appears in the multi-
function display for approximately five sec-
onds, and the corresponding speedometer 
segments from the selected speed to the 
vehicle maximum speed are illuminated.
iDepressing the accelerator pedal does 
not deactivate cruise control. After 
brief acceleration (e.g. for passing), 
cruise control will resume the last 
speed set.
iE 320, E 500:
When you use the cruise control lever 
to decelerate, the brake system will au-
tomatically brake the vehicle if the en-
gine’s braking power does not brake 
the vehicle sufficiently.
E 320 4MATIC, E 500 4MATIC:
When you use the cruise control lever 
to decelerate, the transmission will au-
tomatically downshift if the engine’s 
braking power does not brake the vehi-
cle sufficiently.
Warning!
G
The speed stored in memory should only be 
set again if prevailing road conditions per-
mit. Possible acceleration or deceleration 
differences arising from returning to preset 
speed could cause an accident and/or seri-
ous injury to you and others.